IZW-Berlin : Forschung - FG 2 (Evolutionsgenetik)

Research Group 2: Evolutionary Genetics

Prof. Dr. Simone Sommer

Evolutionary Genetics
Leibniz Institute for Zoo and Wildlife Research (IZW), Berlin
Evolutionary Ecology
Institute of Biochemistry and Biology
University of Potsdam, Potsdam

Research fields:

My research centres on evolutionary ecology and genetics in natural populations, emphasising the relationship between Behavioural Ecology, Population Biology and Evolutionary & Conservation Genetics. We combine ecological field work with lab analyses (parasite screening, genetics) in a wide range of different mammalian taxa (e.g. rodents, marsupials, lemurs, bats, lagomorphs and carnivores) and have ongoing projects in Africa, Central- and South America, Asia, Australia and Europe. We use genetic as well as genomic approaches to investigate the importance of neutral (microsats, SNPs) and adaptive genetic diversity (MHC, major histocompatibility complex) in evolutionary relevant and adaptive processes, as well as the associated fitness consequences. Specifically, we currently focus on

*	Consequences of anthropogenic impact (fragmentation, degradation, climatic changes) and associated ecological changes on behaviour, metapopulation ecology and genetics, microhabitat use and pathogen loads.
*	The role of structural MHC variability and expression pattern of innate and adaptive immune relevant genes in host-pathogen interactions, disease resistance and population fitness.
*	Mechanisms and benefits of female choice and their consequences on fitness, genetic structure and gene flow.
*	Olfactory choice of partners – immune system, MHC-linked olfactory receptors and their adaptive importance for the level of health in mammals.

Education and scientific career:

since 2007	Professor for Evolutionary Ecology, University of Potsdam.
08. Nov 2006	Venia legendi for Evolutionary Ecology, University of Potsdam.
since 2006	Head of Evolutionary Genetics, Leibniz-Institute for Zoo- and Wildlife Research (IZW), Berlin.
2005	Scientific Fellow (German-South African Scientific Exchange Program), University of Stellenbosch/South Africa, Dept Genomics & Conservation Biology (Prof Dr T Robinson, Prof Dr C Matthee and Dr S Matthee), Research topics: Evolutionary Genetics, gastrointestinal parasites of South African mammals.
2004-2006	Assistant Professor (Wissenschaftliche Mitarbeiterin), Animal Ecology and Conservation, University of Hamburg (Prof JU Ganzhorn).
03. Feb 2004	Venia legendi for Zoology and Animal Conservation, University of Hamburg.
28. Jan 2004	Habilitation in Zoology, University of Hamburg, Cumulative thesis: ‘Consequences of social systems and anthropogenic impacts on the population ecology and genetics of animal populations’.
since 2002	Project coordinator and principal investigator, BMBF BIOCAPSP, ‘Consequences of habitat fragmentation on the genetic diversity, parasite resistance, and population ecology of mammals: Biodiversity conservation in fragmented landscapes at the Atlantic Plateau of São Paulo (Brazil) (BioCAPSP)’, funded by the German Federal Ministry of Education and Research (BMBF) `Science and Technology for the Mata Atlântica/Brazil’.
2001	Project coordinator and principal investigator, IUCN Species Survival Commission (world leading conservation agency). ‘Individual based modelling on the effects of anthropogenic impact on the population dynamics of endangered mammal species to define priorities for biodiversity conservation in Madagascar’.
1998-2004	Assistant Professor (Hochschulassistentin, C1), Animal Ecology and Conservation, University of Hamburg (Prof JU Ganzhorn).
1998	Postdoctoral fellow (DAAD), Washington University, St. Louis/USA (Prof Dr A Templeton) and Northwestern University, Chicago (Prof Dr A Yoder).
1995-1997	PhD student, Eberhard-Karls-Universität Tübingen and Max-Planck-Institute for Biology in Tübingen, Dept. Immunogenetics, PhD Thesis: ‘Population ecology and –genetics of Hypogeomys antimena, an endemic rodent of the dry deciduous forest in western Madagascar’.
1995-1997	PhD Fellowship, Landesgraduiertenstipendium Baden-Württemberg.
1995-1997	Teaching and Research Assistant, Institute for Behavioural Physiology, Eberhard-Karls-Universität Tübingen.
1995-1996	Fellow of the German-American Academic Council (Deutsch-Amerikanisches Akademisches Konzil (DAAK), 1. German-American Summer Institute in Ann Arbor/USA und Bielefeld/Germany.
1992-1993	Scientific Fellow, World Wildlife Fund (WWF) Madagascar.
1991	Volunteer for a Nature Conservancy Organisation (Aktionsgemeinschaft Artenschutz): Conservation of the marine sea turtle (Caretta caretta) and the coastal soft-shell turtle (Trionyx triunguis) in Dalyan and Patara (Turkey).
1990-1994	Teaching and Research Assistant, Zoological Institute, Eberhard-Karls-Universität Tübingen.
1990-1994	Study of Biology, Eberhard-Karls-Universität Tübingen, Diploma thesis: ‘Ecology and Social Structure of Hypogeomys antimena, an endemic rodent of the dry deciduous forest in western Madagascar’.
1987-1989	Study of Biology, Ruprecht-Karls-Universität Heidelberg.

Academic honours and awards:

1998	Fritz Frank Award of the German Society for Mammalogy.
2000	Hamburg: “summa cum laude“-Award of the Senat of the City of Hamburg.
2003	Invitation as outstanding Junior Scientist, 92. Dahlem Conference ‘Attachment and Bonding: a new Synthesis’, Berlin.
2007	Invitation as outstanding German Scientist in Natural Sciences, 13. German-American Frontiers of Science Symposium (GAFOS), Irvine California/USA by the Alexander von Humboldt-Foundation and the National Academy of Sciences, USA.

	> 70 invited talks, including >10 plenary talks at International Scientific Conferences.

Professional services for funding agencies and scientific societies:

since 2010	Invited Associate Member of the Centre of Infection Biology and Immunology (ZIBI, Berlin).
since 2009	Scientific Board Member of the Society for Tropical Ecology (GTÖ).
since 2009	Associate Editor: Conservation Genetics..
since 2009	Editorial Board: Ecotropica.
since 2009	Scientific Board Member of the Berliner Centre for Genomics in Biodiversity Research
since 2008	Referee Board Member of the German Science Foundation (DFG-Fachkollegiatin) for Zoology (Evolution, Ecology, Biodiversity and Anthropology).

Ad-hoc Referee for Journals:

Acta Theoretica, Animal Conservation, Basic and Applied Ecology, Behavioural Ecology and Sociobiology, Biological Conservation, Biological Journal of the Linnean Society, BMC Biology, BMC Ecology, BMC Evolutionary Biology, BMC Genetics, BMC Genomics, BMC Research Notes, Conservation Genetics, Ecography, Ecotropica, Endangered Species Research, Gene, Heredity, International Journal of Primatology, Mammalian Biology, Molecular Ecology, Oecologia, Oryx, Philosophical Transactions B, PloS One, Proc R Soc London B, Science, Springer-Verlag, Zoological Science, Zoological Studies.

Ad-hoc Referee for Granting agencies:

German Science Foundation (DFG), German Federal Ministry of Education and Research (BMBF), Umweltforschung Baden-Württemberg (BWPlus)/Germany, German Academic Exchange Service (DAAD), Fond zur Förderung der wissenschaftlichen Forschung (FWF)/Austria, National Research Foundation (NRF)/South Africa, National Science Foundation (NSF)/USA, Sigma Delta Epsilon Fellowships/USA, Graduate Women in Science/USA, National Environmental Research Council (NERC)/UK, Alexander von Humboldt Foundation, Böll-Foundation, Chevening Scholarship/UK, Claude Leon Foundation/South Africa, Schering-Foundation, Studienstiftung des Deutschen Volkes, Universitätsstiftung Hamburg, Tinbergen-Award.

Conference organisation (past 5 years):

2010	Organisation of a symposium on ‘Bat Immunology and Immunogenetics’ at the ‘2nd International Berlin Bat Conference: Bat Biology and Infectious Diseases’, Berlin.
2009	Head of the Organisation Committee of the ‘7th International Conference on Physiology, Behaviour and Genetics of Wildlife Animals’, Berlin (> 350 participants, > 35 countries).
2009	Organisation of the symposium ‘Evolutionary Approaches in Conservation: from Genetics to Genomics’ at the ‘7th International Conference on Physiology, Behaviour and Genetics of Wildlife Animals’, Berlin..
2007	Co-Organisation of the ‘6th International Conference on Physiology, Behaviour and Genetics of Wildlife Animals’, Berlin.
2007	Organisation of the symposium ‘Conservation genetics: The role of immune genes and inbreeding on fitness traits’ at the ‘6th International Conference on Physiology, Behaviour and Genetics of Wildlife Animals’, Berlin.

Major funding and grants (past 5 years):

*	BMBF (01LB 0202, 01LB 0202B),, ‘Consequences of habitat fragmentation on the genetic diversity, parasite resistance, and population ecology of mammals: Biodiversity conservation in fragmented landscapes at the Atlantic Plateau of São Paulo (Brazil): 986.000 €.
*	DFG (So 428/4-1, 4-2), Effects of social system, mate choice, parasite load and MHC-constitution on the individual fitness of two sympactric Malagasy lemur species: 172.000 €.
*	DFG (GA 342/14-1), ’Ecological, physiological and genetic basis leading to the differentiation of mouse lemurs in South Madagascar’: 22.500€ (own share).
*	Volkswagen Foundation, Graduate school ’Evolution across scales’ with colleagues of the University of Potsdam: 300.000€.
*	DFG Priority Program ‘Host-Parasite Coevolution – Rapid reciprocal adaptation and its genetic basis’ (SPP 1399, DFG SO 428/7-1), ‘Host adaptations at the molecular and transcriptional level driven by a fast evolving pathogen, the rabbit haemorrhagic disease virus (RHDV) raging in European rabbits (Oryctolagus cuniculus): 220.000€.
*	Pakt für Innovation und Forschung, ’Olfactory choice of partners – immune system, smell receptors and their adaptive importance for the level of health in mammals’: 1.100.000€.

Selected publications in peer-reviewed journals and book contributions:

Submitted:

Castro-Prieto A, Wachter B, Sommer S. Cheetah paradigm revisited: MHC diversity in the largest free-ranging population.

Froeschke G, Sommer S. Evidence for parasite mediated contemporary selection on the MHC class II DRB genes in the Striped Mouse (Rhabdomys pumilio) along a climatic gradient in southern Africa.

Froeschke G, Sommer S. Insights into the complex associations between MHC class II DRB polymorphism and multiple gastrointestinal parasite infestations in the Striped Mouse (Rhabdomys pumilio) in southern Africa.

Ploshnitsa A, Goltsman M, Kennedy L, MacDonald D, Sommer S. Impact of historical founder effects and a recent bottleneck on MHC variability in Commander Arctic foxes (Alopex lagopus).

Puettker T, Bueno A, Barros C, Sommer S, Pardini R. Underlying demographic mechanisms of context-dependent patch area effects on species abundance.

Schad J, Dechmann D, Voigt C, Sommer S. MHC class II DRB diversity, selection pattern and population structure in a neotropical bat species, Noctilio albiventris.

Chronological order:

Ganzhorn JU, Hapke A, Lahann P, Raharivololona BM, Ramanamanjato J-B, Refaly E, Schmid J, Schad J & Sommer S (2010). Population genetics, parasitism and long-term population dynamics of Microcebus murinus in littoral forest fragments of south-eastern Madagascar. In: Leaping ahead: Advances in Prosimian Biology (Masters J, Gamba M & Génin F, eds). Springer, Heidelberg. In press (accepted 01.11.2009).

Meyer-Lucht Y, Otten C, Püttker T, Pardini R, Metzger JP, Sommer S (2010) Variety matters: adaptive genetic diversity and parasite load in two mouse opossums from the Brazilian Atlantic Forest differing in their sensitivity to habitat fragmentation. Conservation Genetics, in press (accepted 21.04.2010).

Schwensow N, Dausmann K, Eberle M, Fietz J, Sommer S (2010). Functional associations of similar MHC alleles and shared parasite species in two sympatric lemurs. Infection, Genetics and Evolution, in press (accepted 26.03.2010).

Schwensow N, Eberle M, Sommer S (2010). Are there ubiquitous parasite-driven MHC selection mechanisms in gray mouse lemurs? International Journal of Primatology, in press (accepted 09.10.2009).

Sommer S, Taubert R, Schmidt A (2010). Isolation of new microsatellite markers using genome screening and restriction-ligation for Apodemus flavicollis. Added to Permanent Genetic Resources Database (1 February 2010–31 March 2010). Molecular Ecology Resources, in press (accepted 07.02.2010).

Sommer S, Taubert R, Schmidt A, Axtner J, Lieckfeldt D (2010) Development of polymorphic microsatellite marker for Delomys sublineatus, an endemic rodent of the Brazilian Atlantic rain forest by a time and cost-efficient enrichment protocol using genomic DNAs of unrelated organisms for cross-hybridization. Molecular Ecology Resources, in press (accepted 15.05.2010).

Weyrich A, Axtner J, Sommer S (2010). Selection and validation of reference genes for real-time RT-PCR studies in the non-model species Delomys sublineatus, an endemic Brazilian rodent. Biochemical and Biophysical Research Communications, doi:10.1016/j.bbrc.2009.12.173 (accepted 30.12.2009).

Axtner J, Sommer S (2009). Validation of internal reference genes for quantitative real-time PCR in a non-model organism, the yellow-necked mouse, Apodemus flavicollis. BMC Research Notes, 2:264.

Froeschke G, Harf R, Sommer S, Matthee S (2009). Effects of precipitation on parasite burden along a climatic gradient in southern Africa – implications for possible shifts in disease patterns due to global changes. Oikos, in press (accepted 09.10.2009). (SS and MS contributed equally to the study).

Lenz TL, Wells K, Pfeiffer M, Sommer S (2009). Divergent MHC IIB allele repertoire increases parasite resistance and body condition in the Long-tailed giant rat (Leopoldamys sabanus). BMC Evolutionary Biology, 9, 269.

Meyer-Lucht Y, Sommer S (2009). Number of MHC alleles is related to parasite loads in natural populations of yellow necked mice (Apodemus flavicollis). Evolutionary Ecology Research, 11(7), 1085-1097.

Sommer S, Schmidt A, Fernandez F, Püttker T, Pardini R (2009). Development and characterization of microsatellite loci in the marsupial Marmosops incanus (Lund, 1840) of the Brazilian Atlantic rain forest using genome screening and restriction ligation. Added to Permanent Genetic Resources Database (1 May 2009–31 July 2009). Molecular Ecology Resources, 9, 1460-1466.

Ganzhorn JU, Andrianasolo T, Andrianjazalahatra T, Donati G, Fietz J, Lahann P, Norscia I, Rakotondranary J, Rakotondratsima BM, Ralison J, Ramarokoto REAF, Randriamanga S, Rasarimanana S, Rakotosamimanana B, Ramanamanjato JB, Randria G, Rasolofoharivelo MT, Razanahoera-Rakotomalala M, Schmid J, Sommer S (2008). Lemurs in evergreen littoral forest fragments. In: Biodiversity, Ecology and Conservation of Littoral Ecosystems in Southeastern Madagascar (Ganzhorn JU, Goodman SM & Vincelette M, eds). Smithsonian Institution, Washington DC, USA. 223-236.

Meyer-Lucht Y, Otten C, Püttker T, Sommer S (2008). Selection, diversity and evolutionary patterns of the MHC class II DAB in free-ranging Neotropical marsupials. BMC Genetics, 9, 39.

Püttker T, Meyer-Lucht Y, Sommer S (2008). Effect of fragmentation on parasite burden (Nematodes) of generalist and specialist small mammal species in secondary forest fragments of the coastal Atlantic Rain Forest, Brazil. Ecological Research, 23, 207-215.

Püttker T, Meyer-Lucht Y, Sommer S (2008). Fragmentation effects on population density of three rodent species in secondary Atlantic Rainforest, Brazil. Studies on Neotropical Fauna and Environment, 43(1), 11-18.

Püttker T, Pardini R, Meyer-Lucht Y, Sommer S (2008). Responses of five small mammal species to micro-scale variations in vegetation structure in secondary Atlantic Forest remnants, Brazil. BMC Ecology, 8, 9.

Schwensow N, Eberle M, Sommer S (2008). Compatibility counts: MHC-associated mate choice in a wild promiscuous primate. Proceedings of the Royal Society London B, 275, 555-564.

Schwensow N, Fietz J, Dausmann K and Sommer S (2008). MHC-dependent mating strategies and the importance of overall genetic diversity in a pair-living primate. Evolutionary Ecology, 22, 617-636.

Sommer S (2008). Forest fragmentation effects on functional genes: immune gene variability (MHC) of Microcebus murinus and Rattus rattus in the Mandena forest. In: Biodiversity, Ecology and Conservation of Littoral Ecosystems in Southeastern Madagascar (Ganzhorn JU, Goodman SM & Vincelette M, eds). Smithsonian Institution, Washington DC, USA. 259-272.

Axtner J, Sommer S (2007). Gene duplication, allelic diversity, selection processes and adaptive value of MHC class II DRB-genes of the bank vole, Clethrionomys glareolus. Immunogenetics, 59(5), 417-426.

Schwensow N, Fietz J, Dausmann K and Sommer S (2007). Neutral versus adaptive genetic variation in parasite resistance: importance of MHC-supertypes in a free-ranging primate. Heredity, 99, 265 – 277.

Püttker T, Meyer-Lucht Y, Sommer S (2006). Movement distances of five rodent and two marsupial species in forest fragments of the coastal Atlantic Rainforest, Brazil. Ecotropica, 12(2), 131-139.

Froeschke G, Sommer S (2005). MHC Class II DRB constitution and parasite load in the striped mouse, Rhabdomys pumilio, in the Southern Kalahari. Molecular Biology and Evolution, 22(5), 1254-1259.

Harf R, Froeschke G, Sommer S (2005). Effects of different land use on the parasite burden and genetic constitution of two rodent species (Gerbillurus paeba, Rhabdomys pumilio) in the Southern Kalahari Desert. In: African Biodiversity – molecules, organisms, ecosystems (Huber BA, Sinclair BJ & Lampe K-H, eds). Springer Verlag, 351-361.

Harf R, Sommer S (2005). Association between MHC Class II DRB alleles and parasite load in the hairy-footed gerbil, Gerbillurus paeba, in the Southern Kalahari. Molecular Ecology, 14, 85-91.

Hingston M, Goodman SM, Ganzhorn JU, Sommer S (2005). Reconstruction of the colonization of southern Madagascar by introduced Rattus rattus. Journal of Biogeography, 32, 1549-1559.

Meyer-Lucht Y, Sommer S (2005). MHC diversity and the association to nematode parasitism in the yellow necked mouse (Apodemus flavicollis). Molecular Ecology, 14, 2233-2243.

Pederson CA, Ahnert L, Anzenberger G, Belsky J, Draper P, Fleming AS, Grossman K, Sachser N, Sommer S, Tietze DP, Young LJ (2005). Beyond infant attachment – The origins of bonding in later life. In: Attachment and Bonding: A New Synthesis (Carter CS, Ahnert L, Grossmann KE, Hrdy SB, Lamb ME, Porges SW & Sachser N, eds). Dahlem Workshop Report 92. Cambridge, MA. The MIT Press, 385-427.

Schad J, Ganzhorn JU, Sommer S (2005). MHC constitution and parasite burden in the Malagasy mouse lemur, Microcebus murinus. Evolution, 59(2), 439-450.

Sommer S (2005) MHC-dependent mate choice in a monogamous rodent. Behavioural Ecology and Sociobiology, 58, 181-189.

Sommer S (2005) Invited Review: The importance of immune gene variability (MHC) in evolutionary ecology and conservation. Frontiers of Zoology, 2:16.

Musolf, K, Meyer-Lucht Y, Sommer S (2004). Evolution of MHC-DRB class II polymorphism in the genus Apodemus and a comparison of DRB sequences within the family Muridae (Mammalia: Rodentia). Immunogenetics, 56(6), 420-426.

Schad J, Sommer S, Ganzhorn JU (2004). MHC variability of a small lemur in the littoral forest fragments of southeastern Madagascar. Conservation Genetics, 5(3), 299-309.

Sommer S (2003). Effects of habitat fragmentation and changes of dispersal behaviour after a recent population decline on the genetic variability of non-coding and coding DNA of a monogamous rodent. Molecular Ecology, 12, 2845-2851.

Sommer S (2003). Social and reproductive monogamy in rodents: the case of the Malagasy giant jumping rat (Hypogeomys antimena). In: Monogamy: Partnerships in Birds, Humans and other Mammals (Reichard U & Boesch C, eds). Cambridge University Press. 109-124.

Sommer S (2003). Natural history of the Malagasy Giant Jumping Rat, Hypogeomys antimena. In: The Natural History of Madagascar (Goodman S & Benstead J, eds). University of Chicaco Press. 1383-1385.

Sommer S, Schwab D, Ganzhorn JU (2002). MHC diversity of endemic Malagasy rodents in relation to geographic range and social system. Behavioural Ecology and Sociobiology, 51, 214-221.

Sommer S, Seal US (2002). A population and habitat viability assessment (PHVA) for the highly endangered Giant Jumping Rat (Hypogeomys antimena), the largest endemic rodent of Madagascar. In: Evaluation et Plans de Gestion pour la Conservation (CAMP) de la Faune de Madagascar: Lémuriens, autres Mammifères, Reptiles et Amphibians, Poissons d'Eau douce et Evaluation de la Viabilité des Populations et des Habitats de Hypogeomys antimena (Vositse). Conservation Breeding Specialist Group (SSC/IUCN). CBSG, Apple Valley, MN. 101 pp.

Sommer S, Toto Volahy A, Seal US (2002). A population and habitat viability assessment for the highly endangered Giant Jumping Rat (Hypogeomys antimena), the largest extant endemic rodent of Madagascar. Animal Conservation, 5, 263-273.

Ganzhorn JU, Porter P, Lowry II, Schatz GE, Sommer S (2001). Madagascar: one of the world's hottest biodiversity hotspot on its way out. Oryx, 35, 346-348.

Sommer S (2001). Reproductive ecology of the endangered monogamous Malagasy giant jumping rat, Hypogeomys antimena. Mammalian Biology, 66, 111-115.

Sommer S (2000). Sex specific predation on a monogamous rat, Hypogeomys antimena (Muridae: Nesomyinae). Animal Behaviour, 59, 1087-1094.

Sommer S, Hommen U (2000). Modelling the effects of life-history traits and changing ecological conditions on the population dynamics and persistence of the endangered Malagasy giant jumping rat (Hypogeomys antimena). Animal Conservation, 4, 333-343.

Sommer S, Tichy H (1999). MHC-Class II polymorphism and paternity in the monogamous Hypogeomys antimena, the endangered, largest endemic Malagasy rodent. Molecular Ecology, 8, 1259-1272.

Sommer S (1998). Populationsökologie und -genetik von Hypogeomys antimena, einer endemischen Nagerart im Trockenwald Westmadagaskars. Cuvillier Verlag, Göttingen. ISBN 3-89712-055-0.

Sommer S (1997). Monogamy in Hypogeomys antimena, an endemic rodent of the deciduous dry forest in western Madagascar. Journal of Zoology, 241, 301-314.

Ganzhorn JU, Sommer S, Abraham J-P, Ade M, Raharivololona BM, Rakotovao ER, Rakotondrasoa C & Randriamarosoa R (1996). Mammals of the Kirindy Forest with special emphasis on Hypogeomys antimena and the effects of logging on the small mammal fauna. In: Ecology and Economy of a Tropical Dry Forest in Madagascar (Ganzhorn JU & Sorg J-P, eds). Primate Report, 46-1, June 1996, Göttingen. 215-232.

Sommer S (1996). Ecology and social structure of Hypogeomys antimena, an endemic rodent of the deciduous dry forest in western Madagascar. In: Biogeography of Madagascar (Lourenco WR, ed). Editions de l´ORSTOM, Paris. 295-302.

Non-reviewed articles and TV contributions:

Schwensow N, Sommer S (2008). Geheime Wahl bei Lemuren. Verbundjournal der Leibniz-Gemeinschaft, März, 10.

Sommer S (2008) Inseln im Regenwald. Verbundjournal der Leibniz-Gemeinschaft, Juni, 15.

Schwensow N, Sommer S (2007). Partnerwahl hängt von den Genen ab. Verbundjournal der Leibniz-Gemeinschaft, Sept, 7.

Ganzhorn JU, Hennig V, Sommer S (1999). Ökologie und Schutz tropischer Lebensräume. Uni HH, 32, 51-56.

Sommer S (1999). Die Kängururatten von Madagaskar. TV-Sendung über populations-ökologischen Untersuchungen an Hypogeomys antimena in Westmadagaskar: Vox, Tierzeit, Sendetermin: 20.02.1999, 1740 -1820.

Sommer S (1998). Mit Forschern im Gespräch. Das Tier, 5, 68.